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The objective of the speech is to offer a briefly introduction of Uroplakins, a protein family mainly expressed by the differentiated urothelial lining cells “umbrella cells”. Particular attention will be paid to Uroplakin IIIc, a novel divergent uroplakin discovered in the “Molecular Biology of the Healthy and Pathologic Urothelium” IISPV research group led by the Principal Investigator Antonio García-España